Professor of Ecology

Department of Biological Applications and Technology
University of Ioannina
Ioannina 45110, Greece


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Department of Biological Application & Technology

University of Ioannina


Some of the exciting courses that I teach or coordinate at the University of Ioannina and elsewhere.

General Ecology

We carry out introduction to ecology with laboratories in theory, data analysis and fieldwork including an excursion to Zagori.


  1. Populations. Sampling and density, population growth, density dependence, life tables, age-structured populations
  2. Ecological interactions. Interactions (++, +- and –), competition, predation, herbivory, parasitism, symbiosis, coevolution.
  3. Communities and biodiversity. Diversity indices, species abundance relations, types of diversity, SARs, island biogeography.


  1. Population dynamics (PC lab)
  2. Measuring plant biodiversity (UOI campus)
  3. Analysis of diversity data (PC lab)
  4. Field trip to Zagori: self-thinning, life tables and succession
  5. Analysis of field data (PC lab)



  1. Molles, Manuel C. (Jr), Ecology: meaning and application, 2009
  2. Lykakis, S., Ecology, Athanasopopulos-Papadamis Publishers, Athens 1996.
  3. J.M. Halley, Notes for General Ecology, UOI Website.
Applied Ecology

Course Contents
1.    Introduction to Applied Ecology
2.    Ecosystems: Biomes, energy and productivity, trophic chains, decomposition, biogeochemical cycles.
3.    Biodiversity loss: prehistoric extinctions, natural and anthropogenic rates of extinction, IUCN categories, restoration ecology
4.    Resource management: theories of harvesting, social aspects of resource management, tragedy of the commons
5.    Climate change: evidence of global warming, prehistoric and historic changes, causes of change, responses of ecosystems
5.    Ozone hole: importance of ozone at high altitudes and low altitudes, history and politics of the ozone hole
6.    Management of pests and invasive species

Laboratory Exercises
1.    Ecological footprint (students carry out this lab at home)
2.    Population dynamics and genetics of small populations (PC lab)
3.    Estimation of biodiversity and distribution (field experiment)

Course books (in Greek)
1.    Halley, J.M., Notes for Applied Ecology, UOI Website (e-course)
2.    Vokou, D, General Ecology, University Studio Press, Thessaloniki 2009
3.    Tyler-Miller, G., Environmental Sciences, Ion publishers 2004
4.    Lykakis, S., Ecology, Athanasopopulos-Papadamis Publishers, Athens 1996

Field course in Ecology

COORDINATORS – J.M.Halley, K.G.Sotiropoulos
LECTURERS – J.M.Halley, K.G.Sotiropoulos & K. Stara (University of Ioannina), R. Tsiakiris (Forestry Authority), M. Argyropoulou & D. Gioulatos (Aristotle University of Thessaloniki),

This is an elective course. It is for students who want to learn more about ecology in the field.

Comprises 60 hours in total and is worth 5 ECTS. This course involves a series of preparation lectures at the university and then, usually in late May, six days in Zagori, a mountainous area, under guidance for the fieldwork. Students (up to 20) stay at the University research station (PALASE) in Ano Pedina. Work includes measurements and data gathering. The students create a database to establish and analyze the data collected.

Nikos shows insects

PRACTICALS (48 hours)

  1. Techniques for observation of animals in the area. Nocturnal detection methods, acoustics & birdsong analysis, invertebrate trapping.
  2. Natural history. Depending on the instructors, natural history of plants, birds, mammals, reptiles and amphibians, butterflies, other invertebrates.
  3. Ecological data analysis. After each field session, students organize their data and carry out statistical analyses
  4. Biodiversity databases. Basic principles of biodiversity database design. Students in teams create their own dB.

LECTURES (12 hours)

  1. Introduction to field ecology
  2. Review of statistics
  3. Introduction to database design
  4. Field safety and practice
  5. The environment of Zagori


  1. W.J. Sutherland, Ecological Census Techniques (2006) Cambridge University Press.
    Mucking amphib
Greek Summer School in Conservation Ecology

GSS: Biodiversity Theory and Practice.

The Greek Summer schools are held every year at the PALASE field station of the University of Ioannina in the Pindos Mountains of Greece. This series of schools in Conservation Biology began in 2008. The duration of each is 12 days and aimed at graduate and postgraduate students. Applicants with strong quantitative skills are especially welcome. We offer training on issues of biodiversity and advanced statistical methods for conservation biology. Apart from training with modern statistical and other quantitative tools (such as R and database design), participants have the opportunity to experience the stunning biodiversity of the region by working in small teams gathering data from the wide variety of habitat types. Participants can observe how different types of diversity (taxonomic, phylogenetic) have been affected by human culture both negatively, for example after land abandonment, and also in positive ways, such as at sacred natural sites. GSS-2016 is under the auspices of the Hellenic Ecological Society (HELCOS) and Society for Conservation Biology (SCB Europe).

GSS 2014 highlights (image)

The course is worth 6 ECTS and is aimed at postgraduate students; applicants with good quantitative skills are especially welcome. The number of participants is usually limited to 15. As familiarity with R is essential, we offer a special two-day refresher course prior to the start of the school for those wishing to ‘brush up’ on R.

More information can be found at the GSS website.